I also faced this problem and tried a lot of things to get to the root cause of the problem. It was very frustrating as AncestorLevel is not used anywhere in code! and I was not able to find the place in code which is responsible for these errors.System.Windows.Data Error: 4 : Cannot find source for binding with reference 'RelativeSource FindAncestor, AncestorType='System.Windows.Controls.DataGrid', AncestorLevel='1''. BindingExpression:Path=CellsPanelHorizontalOffset; DataItem=null; target element is 'Button' (Name=''); target property is 'Width' (type 'Double') System.Windows.Data Error: 4 : Cannot find source for binding with reference 'RelativeSource FindAncestor, AncestorType='System.Windows.Controls.ItemsControl', AncestorLevel='1''. BindingExpression:Path=HorizontalContentAlignment; DataItem=null; target element is 'ComboBoxItem' (Name=''); target property is 'HorizontalContentAlignment' (type 'HorizontalAlignment') System.Windows.Data Error: 4 : Cannot find source for binding with reference 'RelativeSource FindAncestor, AncestorType='System.Windows.Controls.ItemsControl', AncestorLevel='1''. BindingExpression:Path=VerticalContentAlignment; DataItem=null; target element is 'ComboBoxItem' (Name=''); target property is 'VerticalContentAlignment' (type 'VerticalAlignment')
Even after searching the various forums and articles there was no solution for this problem; but this was a very common issue and cause of this problem as mentioned on various forums:
This is a "known" issue, and happens to all controls that contain dynamically created lists (all item controls i.e. ComboBox, menu, ListBox etc.).Microsoft guys mention here and here that “This error has already been handled internally, so you can just leave it alone.” But, still I wanted some sort of solution so as not to have so many irritating error messages in my output window;
ControlTemplate of items in these controls (specifically MenuItem, ComboBoxItem etc.)
try to find the nearest ItemsControl and bind to the VerticalAlignment and HorizonalAlignment properties and raises this error on not finding the source.
So, I tried one of the workaround provided for this problem i.e. to explicitly set the properties which cause problems like this -
<ComboBoxName="control"><ComboBox.ItemContainerStyle><StyleTargetType="ComboBoxItem"><SetterProperty="HorizontalContentAlignment"Value="Left" /><SetterProperty="VerticalContentAlignment"Value="Center" /></Style></ComboBox.ItemContainerStyle></ComboBox>
This works but it’s very hard task to set these properties across the whole solution for each problematic control(i.e. ListBox, Menu, ContextMenus etc.).
Have a look at this SO question for some other workarounds for this problem - ListBox with Grid as ItemsPanelTemplate produces weird binding errors
Another workaround is to suppress these errors (actually, it seems more appropriate to call them warnings) by setting the data binding source switch level as critical in constructor of the class or a top level window -
I find this one line workaround more appropriate as it works for all such controls (like Menus, ListBox etc.) and that too across the whole project -#if DEBUGSystem.Diagnostics.PresentationTraceSources.DataBindingSource.Switch.Level = System.Diagnostics.SourceLevels.Critical;
REFERENCE : https://weblogs.asp.net/akjoshi/resolving-un-harmful-binding-errors-in-wpf